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- About us
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I am a wildlife biologist conducting research programmes/collaborations in Peru, Ethiopia, China, Indonesia, and in the UK. I am particularly interested in avian ecology and conservation, reintroduction strategies for threatened species, and developing capacity in conservation science. I am a member of the Conservation, Evolution and Behaviour Research Group (CEB) at the Department of Natural Sciences, Manchester Metropolitan University.
Post-graduate Certificate in Academic Practice (2013-2014). Manchester Metropolitan University, UK.
PhD (2002-2007): Department of Environmental & Geographical Sciences, Manchester Metropolitan University, UK. Title: Abundance and ecology of high Andean Polylepis birds in Perú: Implications for habitat management strategies
M.Sc. Conservation Biology (1994-1997): Department of Biological Sciences, Manchester Metropolitan University.
Guest Member of Staff: School of Biology, Newcastle University, UK: (August 2009- April 2011):
Conservation Training Officer: World Pheasant Association, Field Biology Station, Newcastle University, UK (August 2009- April 2011)
Unit Co-ordinator (Undergraduate Biology Network): 6F4Z1005 Tutorial Activities and Field Skills
Unit Co-ordinator (Postgraduate Biology Network): 6F7Z1018 Avian Biology and Conservation
Field Course First Aid Coordinator - Department of Natural Sciences
BSc Biology
BSc Wildlife Biology
BSc Animal Behaviour
MSc Conservation Biology
MSc Bird Conservation
External Examiner for BSc Conservation Biology and MSc Wildlife Conservation: Department of Biological Sciences, University of Chester (2015 - present)
Subject Health Review: Biology Programmes, School of Applied Sciences, University of South Wales (2016)
Avian Ecology and Conservation
Reintroduction Biology
Wildlife Biology
Biodiversity Survey Techniques
Sustainable management of an Ethiopian rangeland for biodiversity and pastoralists. Funded by Darwin Initiative Grant. With: RSPB, BirdLife International, Ethiopian Wildlife and Natural History Society, SOS Sahel, and Coventry University.
Modelling evolution of mate choice in birds: With: Professor Yue-Hua Sun & Dr Nan Lu, Laboratory of Animal Ecology and Conservation Biology, Institute of Zoology, Chinese Academy of Sciences, Beijing, China
Impact of residential and renewable energy development on moorland bird populations, UK. With: Moors for Future Foundation, West Yorkshire Ecology, Bradford Unitary Authority, Calderdale Unitary Authority & Kirklees Unitary Authority (all UK)
Conservation ecology and reintroduction of the Critically Endangered Black-winged Myna in Indonesia. With: Chester Zoo, BirdLife International.
Developing non-invasive bird survey methods for improved long-term avian monitoring programmes. With: Professor Yanyun Zhang, Laboratory for Biodiversity Sciences and Ecological Engineering, College of Life Sciences, Beijing Normal University, China
J. Zhang, Y. Zhang, H. Lloyd, Z. Zhang, D. Li (2021). Rapid reclamation and degradation of suaeda salsa saltmarsh along coastal china’s northern yellow sea. Land. 10(8), pp.835-835.
N. Lyu, MR. Servedio, H. Lloyd, Y-H. Sun (2017). The evolution of post-pairing male mate choice. Evolution. 71(6), pp.1465-1477.
B. Walker, G. Cheshire, H. Lloyd (2007). Peruvian Wildlife A Visitor's Guide to the Central Andes. Bradt Travel Guides.
D. Li, M. Han, H. Lloyd, L. Jin, L. Zhang, et al. J. Yin, D. Wan. (2021). Loss of extra-pair paternity is not associated with decreased paternal investment in mixed-paternity broods or unrelated nestlings in the varied tit, Parus varius. Pakistan Journal of Zoology. 53(6), pp.2105-2116.
C. Yang, J. Wang, N. Lyu, H. Lloyd (2021). Comparison of digital photography and spectrometry for evaluating colour perception in humans and other trichromatic species. Behavioral Ecology and Sociobiology. 75(11),
J. Zhang, Y. Zhang, H. Lloyd, Z. Zhang, D. Li (2021). Rapid reclamation and degradation of suaeda salsa saltmarsh along coastal china’s northern yellow sea. Land. 10(8), pp.835-835.
J. Zhang, H. Zhang, Y. Liu, H. Lloyd, J. Li, et al. Z. Zhang, D. Li. (2021). Saltmarsh vegetation and social environment influence flexible seasonal vigilance strategies for two sympatric migratory curlew species in adjacent coastal habitats. Avian Research. 12(1),
CPE. Bedson, L. Thomas, PM. Wheeler, N. Reid, WE. Harris, et al. H. Lloyd, D. Mallon, R. Preziosi. (2021). Estimating density of mountain hares using distance sampling: a comparison of daylight visual surveys, night-time thermal imaging and camera traps. Wildlife Biology. 2021(3),
C. Xia, H. Lloyd, J. Shi, C. Wei, Y. Zhang (2018). Dawn singing of the Brownish‐flanked Bush Warbler influences dawn chorusing in a bird community. Ethology. 124(6), pp.400-409.
Y. Li, C. Xia, H. Lloyd, D. Li, Y. Zhang (2017). Identification of vocal individuality in male cuckoos using different analytical techniques. Avian Research. 8(1),
N. Lyu, MR. Servedio, H. Lloyd, Y-H. Sun (2017). The evolution of post-pairing male mate choice. Evolution. 71(6), pp.1465-1477.
L. Boye, L. Li, H. Lloyd, X. Canwei, Y. Zhang, et al. G. Zheng. (2016). Comparing post-release survival and habitat use by captive-bred Cabot’s Tragopan (Tragopan caboti) in an experimental test of soft-release reintroduction strategies. Avian Research. 7,
N. Lyu, H. Lloyd, YH. Sun (2015). Delayed plumage maturation in birds and the significance of condition-dependent parental care. Behavioral Ecology and Sociobiology. 69(6), pp.1003-1010.
N. Lyu, H. Lloyd, Y-H. Sun (2015). Delayed plumage maturation in birds and the significance of condition-dependent parental care. Behavioral Ecology and Sociobiology. 69(6), pp.1003-1010.
D. Li, Y. Ding, Y. Yuan, H. Lloyd, Z. Zhang (2014). Female tidal mudflat crabs represent a critical food resource for migratory Red-crowned Cranes in the Yellow River Delta, China. Bird Conservation International. 24(4), pp.416-428.
D. LI, Y. DING, Y. YUAN, HUW. LLOYD, Z. ZHANG (2014). Female tidal mudflat crabs represent a critical food resource for migratory Red-crowned Cranes in the Yellow River Delta, China. Bird Conservation International. 24(4), pp.416-428.
C. Xia, C. Wei, H. Lloyd, J. Liu, Q. Wu, et al. Y. Zhang. (2014). Dawn Singing Intensity of the Male Brownish‐Flanked Bush Warbler: Effects of Territorial Insertions and Number of Neighbors. Ethology. 120(4), pp.324-330.
C. Torres-Sovero, H. Lloyd, DJ. Brightsmith (2014). Clay lick use and other observations of the Blue-headed Macaw (Primolius couloni) in southeastern Peru. ORNITOLOGIA NEOTROPICAL. 25(1), pp.87-94.
D. Li, S. Chen, H. Lloyd, S. Zhu, K. Shan, et al. Z. Zhang. (2013). The importance of artificial habitats to migratory waterbirds within a natural/artificial wetland mosaic, Yellow River Delta, China. Bird Conservation International. 23(2), pp.184-198.
Z. Zhou, Y. Sun, L. Dong, C. Xia, H. Lloyd, et al. Y. Zhang. (2012). Breeding biology of Asian House Martin Delichon dasypus in a high-elevation area. Forktail. pp.62-66.
N. Lu, CX. Jia, H. Lloyd, YH. Sun (2012). Species-specific habitat fragmentation assessment, considering the ecological niche requirements and dispersal capability. Biological Conservation. 152, pp.102-109.
H. Lloyd, S. Sevillano Ríos, SJ. Marsden, A. Valdés-Velásquez (2012). Bird community composition across an Andean tree-line ecotone. Austral Ecology. 37(4), pp.470-478.
N. Lu, Y. Jing, H. Lloyd, YH. Sun (2012). Assessing the distributions and potential risks from climate change for the Sichuan Jay (perisoreus internigrans). Condor. 114(2), pp.365-376.
C. Xia, X. Lin, W. Liu, H. Lloyd, Y. Zhang (2012). Acoustic identification of individuals within large avian populations: a case study of the Brownish-flanked Bush Warbler, South-Central China. PLoS One. 7(8),
CSS. Bernardo, H. Lloyd, F. Olmos, LF. Cancian, M. Galetti (2011). Using post-release monitoring data to optimize avian reintroduction programs: A 2-year case study from the Brazilian Atlantic Rainforest. Animal Conservation. 14(6), pp.676-686.
CSS. BERNARDO, HUW. LLOYD, N. BAYLY, M. GALETTI (2011). Modelling post-release survival of reintroduced Red-billed Curassows Crax blumenbachii. Ibis. 153(3), pp.562-572.
CSS. Bernardo, B. Cresswell, H. Lloyd, R. Azeredo, J. Simpson (2011). Selection of radio transmitter and attachment method for post-release monitoring of captive-bred reintroduced Red-billed Curassow Crax blumenbachii, Brazil. European Journal of Wildlife Research. 57(3), pp.689-694.
M. Wei, H. Lloyd, Y. Zhang (2011). Neighbour–stranger discrimination by Yellow-bellied Tit Parus venustulus: evidence for the “dear-enemy” effect. Journal of Ornithology. 152(2), pp.431-438.
SS. Ríos, H. Lloyd, A. Valdés-Velásquez (2011). Bird species richness, diversity and abundance inPolylepiswoodlands, Huascaran biosphere reserve, Peru. Studies on Neotropical Fauna and Environment. 46(1), pp.69-76.
K. Zhang, N. Yang, Y. Xu, J. Ran, H. Lloyd, et al. B. Yue. (2011). Nesting Behavior of Szechenyi's Monal-Partridge in Treeline Habitats, Pamuling Mountains, China. The Wilson Journal of Ornithology. 123(1), pp.93-96.
D. Li, S. Chen, L. Guan, H. Lloyd, Y. Liu, et al. J. Lv, Z. Zhang. (2011). Patterns of waterbird community composition across a natural and restored wetland landscape mosaic, Yellow River Delta, China. Estuarine, Coastal and Shelf Science. 91(2), pp.325-332.
H. Lloyd, SJ. Marsden (2011). Between-Patch Bird Movements within a High-Andean Polylepis Woodland/Matrix Landscape: Implications for Habitat Restoration. Restoration Ecology. 19(1), pp.74-82.
H. Lloyd (2008). Influence of within-patch habitat quality on high-Andean Polylepis bird abundance. Ibis. 150(4), pp.735-745.
H. Lloyd, SJ. Marsden (2008). Bird community variation across Polylepis woodland fragments and matrix habitats: Implications for biodiversity conservation within a high Andean landscape. Biodiversity and Conservation. 17(11), pp.2645-2660.
H. Lloyd (2008). Foraging ecology of High Andean insectivorous birds in remnant Polylepis forest patches. Wilson Journal of Ornithology. 120(3), pp.531-544.
H. Lloyd (2008). Abundance and patterns of rarity of rarity of Polylepis birds in the Cordillera Vilcanota, southern Perú: Implications for habitat management strategies. Bird Conservation International. 18(2), pp.164-180.
HUW. LLOYD (2004). Habitat and population estimates of some threatened lowland forest bird species in Tambopata, south-east Peru. Bird Conservation International. 14(4), pp.261-277.
H. Lloyd (2003). Population densities of some nocturnal raptor species (Strigidae) in southeastern Peru. Journal of Field Ornithology. 74(4), pp.376-380.
H. Lloyd (2000). Population densities of the Black-faced Cotinga Conioptilon mcilhennyi in south-east Peru. Bird Conservation International. 10(3), pp.277-285.
Member of Editorial Board: Avian Research (Open Access Journal)
Reviewer for journals including: Global Change Biology, Diversity and Distributions, Journal of Applied Ecology, Journal of Animal Ecology, PLoS One, Biological Conservation, Basic and Applied Ecology, Biodiversity and Conservation, Biotropica, Oryx, Ibis, Bird Conservation International and others.